-estris suffix, Latin, origin

From: Joao
Message: 36041
Date: 2005-01-26

What's the origin of Latin suffix -(e)stris? IE or non-IE?
 
terra : terrestris
campus : campestris
ager : agrestis
caelus : caelestis
nemus : nemestris
rupes : rupestris
silva : silvestris
palus : palustris
lacus : lacustris
 
 
 
Joao SL